More Entosthodon

 I was in Mewslade Valley yesterday and noted a 50cm x 20cm patch of E. fascicularis with over 400 densely packed capsules (see photos above - patch on half-covered rock in foreground). I've looked at a few leaves and most have 2 or 3 cells between the costa and the leaf tip, so clearly not of the mouretii type.

I also had a look at the adjacent retaining wall (photos below) and discovered a new colony of E. pulchellus, with numerous small patches scattered along a 100m section of the wall. I counted over 200 capsules, but there were hundreds of non-fruiting plants too. This is possibly the most extensive colony I've personally come across.

One of Britain's least convincing mosses

In 2012 I published a note in Journal of Bryology reporting Entosthodon mouretii as new for Britain, based on 7 records from the southern half of the country.  I initially tried to reduce mouretii to variety status, but one of the journal referees insisted that I could not do that without studying the Type (in Paris), many southern European specimens etc.  I wanted to reduce it to variety status because so many of the British specimens were markedly intermediate, with no obvious discontinuity in the key character states of costa length, leaf acuminateness, and marginal tooth sharpness.  The 7 British records had convincingly excurrent costae, but others had the costa ending 1-5 cells below the apex.

Today I found a substantial fruiting population of E. fascicularis on a roadside bank in Cross Hands Industrial Estate.  Most of its leaves had very long costae, ranging from excurrent by 1 cell to finishing 2 cells below the apex.  It isn't quite extreme enough to be E. mouretii, but it certainly isn't classic short-nerved E. fascicularis.  I am ever more unconvinced at the validity of this taxon as a species, but don't have the time or inclination to shoot it down globally.