Friday, 28 November 2014


In reply to Charles' request for habitat lists - that is something I have been doing to a certain extent during my south Wales recording over the last 15 years.  It's all a bit imprecise, with my habitats and substrates being rather muddled up.  They include various things such as Churchyard, dune slack, tarmac, rocks by stream (and rocks by river), plus virtually every common tree species.  The trees are tabulated in the Pembrokeshire Bryo Flora, against all the regular epiphytes.

Anyway, there are 130,000 bryophyte records from Carms and Pembs in my dataset, 1900 of which have Tarmac as substrate. The following lists the 81 species that I have recorded at least once on Tarmac in SW Wales, plus the number of records of that species.  Rather few species are regular and the 'tail' of one-offs is very long.

Bryum argenteum 306
Bryum dichotomum 298
Didymodon nicholsonii 253
Ceratodon purpureus 116
Cratoneuron filicinum 93
Bryum capillare var. capillare 85
Barbula unguiculata 77
Syntrichia intermedia 72
Syntrichia latifolia 69
Didymodon insulanus 63
Syntrichia ruralis 56
Brachythecium mildeanum 29
Leptodictyum riparium 29
Calliergonella cuspidata 25
Funaria hygrometrica 22
Didymodon luridus 20
Orthotrichum diaphanum 20
Brachythecium plumosum 19
Didymodon sinuosus 19
Schistidium crassipilum 19
Brachythecium rutabulum 17
Tortula truncata 16
Syntrichia ruraliformis 13
Brachythecium albicans 12
Rhynchostegium riparioides 11
Cinclidotus fontinaloides 9
Dialytrichia mucronata 9
Pseudocrossidium hornschuchianum 8
Syntrichia papillosa 8
Barbula convoluta var. commutata 7
Bryum ruderale 7
Philonotis fontana 7
Scleropodium cespitans 7
Tortula muralis var. muralis 7
Barbula convoluta var. convoluta 6
Pohlia wahlenbergii var. wahlenbergii 6
Amblystegium serpens var. serpens 5
Didymodon rigidulus 5
Grimmia pulvinata var. pulvinata 5
Schistidium apocarpum s.s. 5
Bryoerythrophyllum recurvirostrum 4
Bryum capillare 4
Campylopus introflexus 4
Didymodon fallax 4
Encalypta streptocarpa 4
Eurhynchium hians 4
Hypnum cupressiforme 4
Orthotrichum anomalum 4
Orthotrichum cupulatum var. cupulatum 4
Plagiomnium rostratum 4
Brachythecium rivulare 3
Plagiomnium undulatum 3
Schistidium apocarpum 3
Bryum rubens 2
Homalothecium sericeum 2
Hypnum lindbergii 2
Leskea polycarpa 2
Rhizomnium punctatum 2
Rhytidiadelphus squarrosus 2
Scorpiurium circinatum 2
Syntrichia virescens 2
Tortella nitida 2
Amblystegium tenax 1
Barbula convoluta 1
Bryum pseudotriquetrum s.l. 1
Dichodontium sp. 1
Didymodon vinealis 1
Homalothecium lutescens 1
Hygrohypnum luridum var. luridum 1
Hypnum lacunosum var. lacunosum 1
Orthotrichum cupulatum 1
Plagiomnium affine 1
Plagiomnium cuspidatum 1
Rhynchostegium confertum 1
Rhynchostegium murale 1
Rhytidiadelphus triquetrus 1
Schistidium elegantulum 1
Schistidium rivulare 1
Syntrichia laevipila var. laevipila 1
Syntrichia laevipila var. laevipilaeformis 1
Trichostomum crispulum 1


  1. Sam, that is excellent! And, given the eclectic ecologies represented, it is interesting that there isn't a single liverwort there.

  2. I should see if I can find Marchantia polymorpha ruderalis on tarmac in Cardiff. It's really common on flagged pavements but might creep onto tarmac in places...

  3. M.p.r. grows in cracks in weathered tarmac in my street - does that count?
    Great list and reference for future searches - I bet everyone of us looks at the pavements differently tomorrow!

  4. Yes, that counts. What about leafy liverworts then?